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The original model of the viral capsid structure suggested by Watson and Crick, in 1956, viewed the capsid as a curved, two-dimensional “surface crystal” (as opposed to the space-filling 3D crystal), closed in on itself and made of identical subunits — capsid proteins. As each subunit was identical, they interacted in the same way, as in true crystals. But such a model only permitted a maximum of exactly 60 subunits, since the icosahedron has 20 triangular faces, and each triangle can be made of three protein molecules.